as i see your case, i can only suspect what cause the issue although i am no expert on the matter.
sea serpents was originally intended to built with 0.2 layer in mind to short cut as there will be long built time and cost.
i admit there is indeed some geometrical mistake at base of horn as i did't caped and left if opend before i upload the model but you have fixed it with Print studio.
as i see your 0.1 layers setting and the result i suspect your machine's have some troubles with such as nozel cooling capability, bed vibration and Feed stability but not sure as only i can guess it.
i am sorry that i can not help you enough.